The author worked with two main study groups of gorilla in that region. Each contained a fully mature male (silverback), who was the obvious leader of the group, and a younger male of around 9 years of age (blackback), who had not yet assumed full adult morphology.
The second-ranking silverback is also a possible sire in the sixth case, although three other lower-ranking blackback males were also not excluded.
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winter flounder — noun 1. flesh of American flounder; important in the winter • Syn: ↑lemon sole • Hypernyms: ↑flounder • Part Holonyms: ↑blackback flounder, ↑lemon sole, ↑Pseudopleuronectes americanus … Useful english dictionary
